Ecology
The word ecology was coined in 1870 by the
German zoologist Ernst Haeckel from the Greek words oikos (house) and logos
(logic or knowledge) to describe the scientific study of the relationships
among organisms and their environment. Biologists began referring to
themselves as ecologists at the end of the nineteenth century and shortly
thereafter the first ecological societies and journals appeared. The
contextual, historical understanding of organisms as well as the systems
basis of ecology set it apart from the reductionist, experimental approach
prevalent in many other areas of science.
This broad ecological view is gaining significance today as modern
resource-intensive lifestyles consume much of nature's supplies. Although
intuitive ecology has always been a part of some cultures, current
environmental crises make a systematic, scientific understanding of
ecological principles especially important.
For many ecologists the basic structural units of ecological organization
are species and populations. A biological species consists of all the
organisms potentially able to interbreed under natural conditions and to
produce fertile offspring. A population consists of all the members of a
single species occupying a common geographical area at the same time. An
ecological community is composed of a number of populations that live and
interact in a specific region.
This population-community view of ecology is grounded in natural history—the
study of where and how organisms live—and the Darwinian theory of natural
selection and evolution. Proponents of this approach generally view
ecological systems primarily as networks of interacting organisms. Abiotic
forces such as weather, soils, and topography are often regarded as external
factors that influence but are apart from the central living core of the
system.
In the past three decades the emphasis on species, populations, and
communities in ecology has been replaced by a more quantitative,
thermodynamic analysis of the processes through which energy flows and the
cycling of nutrients and toxins are carried out in ecosystems. This
process-functional approach is concerned more with the ecosystem as a whole
than the particular species or populations that make it up. In this
perspective, both the living organisms and the abiotic physical components
of the environment are equal members of the system.
The feeding relationships among different species in a community are a key
to understanding ecosystem function. Who eats whom, where, how, and when
determine how energy and materials move through the system. They also
influence natural selection, evolution, and species adaptation to a
particular set of environmental conditions. Ecosystems are open systems,
insofar as energy and materials flow through them. Nutrients, however, are
often recycled extremely efficiently so that the annual losses to sediments
or through surface water runoff are relatively small in many mature
ecosystems. In undisturbed tropical rain forests, for instance, nearly 100%
of leaves and detritus are decomposed and recycled within a few days after
they fall to the forest floor.
Because of thermodynamic losses every time energy is exchanged between
organisms or converted from one form to another, an external energy source
is an indispensable component of every ecological system. Green plants
capture solar energy through photosynthesis and convert it into energy-rich
organic compounds that are the basis for all other life in the community.